Hacrobia

Hacrobia
	The coccolithophore Gephyrocapsa oceanica
Scientific classification ; (obsolete as its monophyly is doubtful)
Domain:	Eukaryota
(unranked):	Orthokaryotes
(unranked):	Neokaryotes
(unranked):	Bikonta
Superkingdom:	Corticata
Kingdom:	Chromista
Subkingdom:	; Okamoto et al. 2009
Phyla
	Cryptista; Haptista;
Synonyms
	CCTH; Eukaryomonadae;

The cryptomonads-haptophytes assemblage is a proposed but disputed monophyletic grouping^[1] of unicellular eukaryotes that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" referring to Haptophyta, "-cr-" referring to cryptomonads, and "-bia" as a general suffix referring to life);^[2] CCTH (standing for Cryptophyta, Centrohelida, Telonemia and Haptophyta);^[3] and Eukaryomonadae.^[4]

As of February 2012, it is unclear whether this group is monophyletic or not; results of phylogenetic studies are "often dependent on the selection of taxa and gene data set".^[3] Two 2012 studies produced opposite results.^[3]^[5]

^ Sakaguchi M, Takishita K, Matsumoto T, Hashimoto T, Inagaki Y (July 2009). "Tracing back EFL gene evolution in the cryptomonads-haptophytes assemblage: separate origins of EFL genes in haptophytes, photosynthetic cryptomonads, and goniomonads". Gene. 441 (1–2): 126–31. doi:10.1016/j.gene.2008.05.010. PMID 18585873.
^ Okamoto, N.; Chantangsi, C.; Horák, A.; Leander, B.; Keeling, P.; Stajich, J. E. (2009). Stajich, Jason E. (ed.). "Molecular Phylogeny and Description of the Novel Katablepharid Roombia truncata gen. et sp. nov., and Establishment of the Hacrobia Taxon nov". PLOS ONE. 4 (9): e7080. Bibcode:2009PLoSO...4.7080O. doi:10.1371/journal.pone.0007080. PMC 2741603. PMID 19759916.
^ ^a ^b ^c Zhao, Sen; Burki, Fabien; Bråte, Jon; Keeling, Patrick J.; Klaveness, Dag; Shalchian-Tabrizi, Kamran (2012). "Collodictyon—An Ancient Lineage in the Tree of Eukaryotes". Molecular Biology and Evolution. 29 (6): 1557–68. doi:10.1093/molbev/mss001. PMC 3351787. PMID 22319147.
^ "EUKARYOMONADAE". Archived from the original on 2016-03-04. Retrieved 2015-03-01.
^ Burki, F.; Okamoto, N.; Pombert, J.F. & Keeling, P.J. (2012). "The evolutionary history of haptophytes and cryptophytes: phylogenomic evidence for separate origins". Proc. Biol. Sci. 279 (1736): 2246–54. doi:10.1098/rspb.2011.2301. PMC 3321700. PMID 22298847.

[pmid18585873-1] Sakaguchi M, Takishita K, Matsumoto T, Hashimoto T, Inagaki Y (July 2009). "Tracing back EFL gene evolution in the cryptomonads-haptophytes assemblage: separate origins of EFL genes in haptophytes, photosynthetic cryptomonads, and goniomonads". Gene. 441 (1–2): 126–31. doi:10.1016/j.gene.2008.05.010. PMID 18585873.

[okamoto2009-2] Okamoto, N.; Chantangsi, C.; Horák, A.; Leander, B.; Keeling, P.; Stajich, J. E. (2009). Stajich, Jason E. (ed.). "Molecular Phylogeny and Description of the Novel Katablepharid Roombia truncata gen. et sp. nov., and Establishment of the Hacrobia Taxon nov". PLOS ONE. 4 (9): e7080. Bibcode:2009PLoSO...4.7080O. doi:10.1371/journal.pone.0007080. PMC 2741603. PMID 19759916.

[ZhaoBurkBratKeel12-3] Zhao, Sen; Burki, Fabien; Bråte, Jon; Keeling, Patrick J.; Klaveness, Dag; Shalchian-Tabrizi, Kamran (2012). "Collodictyon—An Ancient Lineage in the Tree of Eukaryotes". Molecular Biology and Evolution. 29 (6): 1557–68. doi:10.1093/molbev/mss001. PMC 3351787. PMID 22319147.

[urlEUKARYOMONADAE-4] "EUKARYOMONADAE". Archived from the original on 2016-03-04. Retrieved 2015-03-01.

[Burki2012-5] Burki, F.; Okamoto, N.; Pombert, J.F. & Keeling, P.J. (2012). "The evolutionary history of haptophytes and cryptophytes: phylogenomic evidence for separate origins". Proc. Biol. Sci. 279 (1736): 2246–54. doi:10.1098/rspb.2011.2301. PMC 3321700. PMID 22298847.

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